Grennan, E J., (1999) Lamb growth rate on pasture: Effect of grazing management, sward type and supplementation. Teagasc – End of project reports: Sheep series

No. 3.

Gronvold, J. and Hogh-Schmidt, K. (1989) Factors affecting rain splash dispersal of infective larvae of Ostertagia ostertagi (Trichostrongylidae) from cow pats to the

surroundings. Veterinary Parasitology 31, 57-70.

Gruner, L. and Sauve, C. (1982) The distribution of trichostrongyle infective larvae on pasture and grazing behaviour in calves. Veterinary Parasitology 11, 203-213

Holmes W., (ed) Grass, Its production and utilization. 2nd edition, Blackwell Scientific Publications

Hoste, H., Jackson, F, Athanasiadou, S., Thamsborg, S M and Hoskin, S O. (2006) The effects of tannin-rich plants on parasitic nematodes in ruminants. Trends in

Parasitology 22, No. 6, 253-261

Hutchings, N R., Kyriazakis, I., Anderson, D H., Gordon, I J. and Coop, R L. (1998) Behavioural strategies used by parasitized and non-parasitized to avoid ingestion of

gastro-intestinal nematodes associated with faeces. Animal Science 67, 97-106

Institute of Oragnic Training and Advice (IOTA), Research review – The role and management of herbal pastures for animal health, productivity and product quality.

Jordan, H E., Phillips, W A., Morrison, R D., Doyle, J J. and McKensie, K. (1988) A 3-year study of continuous mixed grazing of cattle and sheep: parasitism of offspring.

International Journal of Parasitology 18, 779-784.

Knapp, S E. (1964) Relationship of different species of forage to the survival and infectivity of Haemonchus contortus in lambs. The Journal of Parasitology 50, 144-


Laca, E A., Ungar, E D., Seligman, N. and Demment, M W. (1992). Effects of sward height and bulk density on bite dimensions of cattle grazing homogenous swards.

Grass and Forage Science 47, 91-102.

Li, G and Kemp, P D. (2005) Forage chicory (Cichorium intybus): A review of its agronomy and animal production. Advances in agronomy 88, 187-222.

Marley, C L., Cook, R., Keatinge, R. Barrett, J. and Lampkin, N H., (2003a) The effect of birdsfoot trefoil (Lotus corniculatus) abd chicory (Cichorium intybus) on

parasite intensities and performance of lambs naturally infected with helminth parasites. Veterrinary Parasitology 112, 147-155

Marley, C L., Cook, R., Barrett, J., Keatinge, R., Lampkin, N H. and McBride, S D. (2003b) The effect of dietary forage on the development and survival of helminth

parasites in ovine faeces. Veterinary Parasitology 118, 93-107

Marley, C L., Fraser, M D., Roberts, J E., Fychan, R. and Jones, R. (2006) Effects of legume forages on ovine gastrointestinal parasite development, migration and

survival. Veterinary Parasitology 138 pp308-317

Marley C L, Cook, R, Barrett, J. Keatinge R and Lampkin N H. (2006a) The effects of birdsfoot trefoil (Lotus corniculatu) and chicory (Cichorium intybus ) when compared with perennial ryegrass (Lolium perenne) on ovine gastrointestinal parasite development, survival and migration. Veterinary Parasitology 138 p280-290.

Marley C L, Fychan R. and Jones R. (2006b) Yield persistency and chemical composition of Lotus species and varieties (birdsfoot trefoil and greater birdsfoot

trefoil) when harvested for silage in the UK. Grass and Forage Science 61, No. 2, 134-145

Marley C L, Fraser, M D., Davies, D A., Rees, M E., Vale, J E. and Forbes A B..(2006c) The effects of birdsfoot trefoil (Lotus corniculatu) and chicory (Cichorium

intybus ) when compared with perennial ryegrass (Lolium perenne) on ovine gastrointestinal parasite development, survival and migration. Veterinary

Parasitology 138 p280-290.

Moss, R A. and Bray, A R. (2006) Effect of sward density and size of faecal deposit on the development and persistence of third-stage Trichostrongylid larvae of sheep.

New Zealand Journal of Agricultural Research 49, 475-481

Moss, R A. and Vlassof, A. (1993). Effect of herbage species on gastro-intestinal roundworm populations and their distribution. New Zealand Journal of Agricultural

Research 36, 371-375

Niezen , J H. Charleston, W A G., Hodgson, J., Mackay, A D and Leathwick, D M. (1996) Controlling internal parasites in grazing ruminants without recourse to

anthelmintics: approaches, experiences and prospects. International Journal of Parasitology 26, 983-992

Niezen , J H. Charleston, W A G., Hodgson, J and Waghorn, T S. (1993) Effect of four grass species on lamb parasitism and growth. Proceedings of the New Zealand

Grassland Association 55, 203-206 

Niezen , J H. Charleston, W A G., Hodgson, J., Miller C M., Waghorn, T S. and Robertson, H A. (1998a) Effect of plant species on the larvae of gastrointestinal

nematodes which parasitise sheep. International Journal of Parasitology 28, 791-803

Niezen , J H. Robertson, H A., Waghorn, G C. and Charleston, W A G. (1998b) Production, faecal egg counts and worm burdens of ewe lambs which grazed six

contrasting forages. Veterinary Parasitology 80, 15-27

Niezen , J H. Robertson, H A., Sidey, A. and Wilson, S R. (2002) The effect of pasture species on parasitism and performance of lambs grazing one of three grasswhite

clover pasture swards. Veterinary Parasitology 105, 303-315

O’Connor L J, Walkden-Brown S W and Kahn L P. (2006). Ecology of the free-living stages of major trichostrongylid parasites of sheep. Veterinary Parasitology 142 1-15

Ramirez-Restepo, C A. and Barry, T N. (2005) Alternative temperate forages containing secondary compounds for improving sustainable productivity in grazing

ruminants. Animal Feed Science and Technology 120 179-201

Scales, G H., Knight, T L. and Saville, D J. (1994) Effect of herbage species and feeding level on internal parasites and production performance of grazing lambs.

New Zealand Journal of Agricultural Research 38, 237-247

Soil Association (2000), Roundworm control in organic sheep systems – technical guide for organic crop and livestock production. 

Stromberg B E. (1997) Environmental factors influencing transmission. Veterinary Parasitology 72, p247-264

Stromberg, B E. and Averbeck, G A. (1999) The role of parasite epidemiology in the management of grazing cattle. Arasitology 29, 33-39

Taylor, M A. (2010) COWS, Control of worms sustainably – sustainable control strategies for cattle

Ungar, E D. and Ravid, N. (1999) Bite horizons and dimensions for cattle grazing herbage to high levels of depletion. Grass and Forage Science 54, 357-364

van Dijk, J. de Louw, M D E. Kalis, L P A. and Morgan, E R. (2009) Ultraviolet light increases mortality of nematode larvae and can explain patterns of larval availability

at pasture. International Journal of Parasitology 39, 1151-1156

van Dijk, J. and Morgan, E R. (2008) The influence of temperature on the development, hatching and survival of Nematodirus battus larvae. Parasitology 135,


Waller, P J. (1997) Sustainable helminth control of ruminants in developing countries. Veterinary Parasitology 71, 195-207

Wormwise - National Worm Management Strategy - A Handbook of Sustainable worm management for livestock farmers. Produced by Meat and Wool New Zealand

Reid JFS and Armour J (1975) Seasonal variation in the gastro-intestinal nematode populations of Scottish hill sheep. Res Vet Sci 18; 307-313.

Cornwell RL (1975) The pattern of infection with gastro-intestinal nematode infection in young fattening lambs at pasture. Res Vet Sci 18; 1-5.

Michel J.F. (1976) The epidemiology and control of some nematode infections in grazing animals. Advances in Parasitology 14, 355-397.

Lancaster MB and Hong C (1988). Observations on the pattern of nematode larval availability on pastures grazed by sheep in Surrey. Vet Rec 122; 235

Learmount J, Taylor M.A., Smith G.C., Morgan C. (2006). A computer model to simulate control of parasitic gastroenteritis in sheep on UK farms. Veterinary Parasitology 142, 213-


Coles GC (2002) Sustainable use of anthelmintics in grazing animals. Vet Rec 151;165-169

Dobson RJ, Besier RB, Barnes EH, Love SCJ, Vizard A, Bell K and LeJambre LF (2001) Principles for the use of macrocyclic lactones to minimise selection for resistance. Aust Vet J 79; 756-761.

Jackson F (1993) Anthelmintic resistance – the state of play. Br Vet J 149;123-138

Leathwick DM, Pomroy WE and Heath ACG (2001) Anthelmintic resistance in New Zealand. NZ Vet J 49; 227-235.

Leathwick DM, Vlassoff A and Barlow ND (1995) A model for nematodiasis in New Zealand Lambs: the effect of drenching regime and grazing management on the development of anthelmintic resistance. Int J Parasitol 25; 1479-1490.

Sargison N (2000) Strategies to limit the development of anthelmintic resistance in sheep flocks. UK Vet 5; 54-57.

Sargison ND and Scott PR (2003) Survey of sheep nematode parasite control methods in south-east Scotland. Vet Rec 152; 51-52.

Sargison ND, Jackson F, Bartley DJ and Moir AC (2005). Failure of moxidectin to control benzimidazole, levamisole and ivermectin resistant Teladorsagia ciumcincta in a sheep flock Vet Rec 156 105-109

Taylor MA, Hunt KR and Goodyear KL (2002) Anthelmintic resistance detection methods. Vet Parasitol 103; 183-194.

Mitchell GBB (2007) Liver fluke. In: Diseases of sheep. ID Aitken (Ed), 4th ed. Blackwell Sciences Ltd, Oxford; 195-204.

Fairweather I and Boray JC (1999) Fasciolicides: Efficacy, actions, resistance and its management. Vet J 158; 81-112.

Target Worm Control for Better Returns. No 8 in BRP Manuals Series. (2008)

Colditz I.G., Le Jambre L.F. and Hosse R. (2002). Use of lectin binding characteristics to identify gastrointestinal parasite eggs in faeces. Veterinary Parasitology 105, 219-227

Colditz I.G. and Le Jambre L.F (2008) Development of a faecal occult blood test to determine the severity of Haemonchus contortus infections in sheep. Veterinary Parasitology 153, 93-99

SCOPS Publications:

All of the following (including this publication) can be found at in pdf format:

Ectoparasite Control in Sheep. Proceedings of a Workshop February 2005

Proceedings of the Technical Review of SCOPS Recommendations January 2009


• 10 things You Should Know (if you want to keep farming sheep)

• New Worm Control Strategies for Sheep. Taking the First Steps

• Correct Drenching Technique

• Quarantine treatments

• Sheep Scab Diagnosis and Control

• Blowfly Control

• Know Your Anthelmintics (2009)